Exploring the effects of habitat management on grassland biodiversity: A case study from northern Serbia

66. Chabuz W, Kulik M, Sawicka-Zugaj W, Żo´łkiewski P, Warda M, Pluta M, et al. Impact of the type of use of permanent grasslands areas in mountainous regions on the floristic diversity of habitats and animal welfare. Glob Ecol Conserv. 2019; 19: e00629. https://doi.org/10.1016/j.gecco.2019.e00629 67. So¨ derstro¨ m B, Svensson B, Vessby K, Glimska¨ r A. Plants, insects and birds in semi-natural pastures in relation to local habitat and landscape factors. Biodivers Conserv. 2001; 10: 1839–1863. https://doi. org/10.1023/A:1013153427422 68. Morandin LA, Kremen C. Hedgerow restoration promotes pollinator populations and exports native bees to adjacent fields. Ecol Appl. 2013; 23(4), 829–839. https://doi.org/10.1890/12-1051.1 PMID: 23865233 69. Donkersley P. Tress for bees. Agric Ecosyst Environ 2019; 270–271: 79–83. https://doi.org/10.1016/j. agee.2018.10.024 70. Krahulec F, Skalova H, Herben T, Hadincova´ V, Wildova´ R, Pecha´ čkova´ S. Vegetation changes follow- ing sheep grazing in abandoned mountain meadows. Appl Veg Sci. 2001; 4: 97–102. https://doi.org/10. 1111/j.1654-109X.2001.tb00239.x. 71. Hellstro¨m K, Huhta AP, Rautio P, Tuomi J, Oksanen J, Laine K. Use of sheep grazing in the restoration of semi-natural meadows in northern Finland. Appl Veg Sci. 2003; 6: 45–52. https://doi.org/10.1111/j. 1654-109X.2003.tb00563.x 72. Danner N, Molitor AM, Schiele S, Ha¨ rtel S, Steffan-Dewenter I. Season and landscape composition affect pollen foraging distances and habitat use of honey bees. Ecol Appl. 2016; 26(6):1920–1929. https://doi.org/10.1890/15-1840.1 PMID: 27755712.

73. Carvell C. Habitat use and conservation of bumblebees (Bombus spp.) under different grassland man- agement regimes. Biol Conserv. 2002; 103: 33–49. https://doi.org/10.1016/S0006-3207(01)00114-8 74. Holzschuh A, Dainese M, Gonza´lez-Varo JP, Mudri-Stojnić S, Riedinger V, Rundlo¨f M, et al. Mass-flow- ering crops dilute pollinator abundance in agricultural landscapes across Europe. Ecol Lett. 2016; 19: 1228–1236. https://doi.org/10.1111/ele.12657 PMID: 27531385 75. Milinski L, Radisˇić D, Arok M, Nikolić T. Foraging habitat characteristics influence the nest- box occupancy and breeding parameters of European roller (Coracias garrulus) in Serbia. Arch Biol Sci. 2022; 74(3): 251–262. https://doi.org/10.2298/ABS220404024M 76. Cousins SAO, Eriksson O. After the hotspots are gone: Land use history and grassland plant species diversity in a strongly transformed agricultural landscape. Appl Veg Sci. 2008; 11: 365–374. https://doi. org/10.3170/2008-7-18480 77. Lindborg R, Eriksson O. Effects of restoration on plant species richness and composition in Scandina- vian seminatural grasslands. Restor Ecol 2004; 12: 318–326. https://doi.org/10.1111/j.1061-2971. 2004.00334.x 78. Helm A, Hanski I, Pa¨rtel M. Slow response of plant species richness to habitat loss and fragmentation. Ecol Lett. 2006; 9: 72–77. https://doi.org/10.1111/j.1461-0248.2005.00841.x. PMID: 16958870 79. Butaye J, Adriaens D, Honnay O. (2005). Conservation and restoration of calcareous grasslands: a con- cise review of the effects of fragmentation and management on lantspecies. Biotechnol Agron Soc Envi-

ron. 2005; 9: 111–118. 80. Piqueray J, Mahy G. Revue bibliographique sur la restauration des pelouses calcicoles en Europe: con- traintes rencontre´es et solutions propose´es. Biotechnol Agron Soc Envir. 2010; 14: 471–484. 81. Rysiaka A, ChabuzbW, Sawicka-Zugaj W, Zdulskib J, Grzywaczewskic G, Kulik M. Comparative impacts of grazing and mowing on the floristics of grasslands in the buffer zone of Polesie National Park, eastern Poland. Glob Ecol Conserv. 2021; 27: Article e01612. https://doi.org/10.1016/j.gecco. 2021.e01612 82. Kapa´ s RE, Plue J, Kimberley A, Cousins SAO. Grazing livestock increases both vegetation and seed bank diversity in remnant and restored grasslands. J Veg Sci. 2020; 31; 1053–1065. https://doi.org/10. 1111/jvs.12956. 83. Littlewood NA, Stewart AJA, Woodcock BA. Science into practice: how can fundamental science con- tribute to better management of grasslands for invertebrates? Insect Conserv Divers. 2012; 5: 1–8. https://doi.org/10.1111/j.1752-4598.2011.00174.x 84. Adriaens D, Honnay O, Hermy M. No evidence of a plant extinction debt in highly fragmented calcareous grasslands in Belgium. Biol Conserv. 2006; 133: 212–224. https://doi.org/10.1016/j.biocon.2006.06.006 85. Honnay O, Coart E, Butaye J, Adriaens D, Van Glabeke S, Roldan-Ruiz I. Low impact of present and historical landscape configuration on the genetics of fragmented Anthyllis vulneraria populations. Biol Conserv. 2006; 127: 411–419. https://doi.org/10.1016/j.biocon.2005.09.006 86. Rupprecht D, Gilhaus K, Ho¨lzel N. Effects of year-round grazing on the vegetation of nutrient-poor

grassand heathlands—Evidence from a large-scale survey. Agric Ecosyst Environ. 2016; 234: 16–22. https://doi.org/10.1016/j.agee.2016.02.015 PLOS ONE Effects of habitat management on biodiversity PLOS ONE | https://doi.org/10.1371/journal.pone.0301391 March 28, 2024 19 / 19