Grazing and light modify Silene latifolia responses to nutrients and future climate

80. Bates D, Maechler M, Bolker B, Walker S. Fitting Linear Mixed-Effects Models Using lme4. Journal of Statistical Software. 2015; 67:1–48. 81. Fox J, Weisberg S. An {R} Companion to Applied Regression. Thousand Oaks CA: Sage; 2019. 82. Wickham H, Averick M, Bryan J, Chang W, McGowan L, Franc¸ois R, et al. Welcome to the Tidyverse. Journal of Open Source Software. 2019; 4(43):1686. 83. Wickham H. ggplot2: Elegant Graphics for Data Analyis. New York: Springer-Verlag; 2016. 84. Pedersen TL. patchwork: The Composer of Plots. 1.1.1 ed 2020. 85. R Core Team. R: A language and environment for statistical computing. 4.0.3 ed. Vienna, Austria: R Foundation for Statistical Computing; 2020. 86. Valladares F, Niinemets U. Shade tolerance, a key plant feature of complex nature and consequences. Annual Review of Ecology, Evolution, and Systematics. 2008; 39(1):237–57. 87. Kahl SM, Lenhard M, Joshi J, Teller B. Compensatory mechanisms to climate change in the widely dis- tributed species Silene vulgaris. Journal of Ecology. 2019; 107(4):1918–30. 88. Chollet S, Rambal S, Fayolle A, Hubert D, Foulquie D, Garnier E. Combined effects of climate, resource availability, and plant traits on biomass produced in a Mediterranean rangeland. Ecology. 2014; 95(3):737–48. https://doi.org/10.1890/13-0751.1 PMID: 24804457 89. Le Bagousse-Pinguet Y, Gross N, Maestre FT, Maire V, Bello F, Fonseca CR, et al. Testing the envi- ronmental filtering concept in global drylands. Journal of Ecology. 2017; 105(4):1058–69. https://doi. org/10.1111/1365-2745.12735 PMID: 28642625

90. Schroeder-Georgi T, Wirth C, Nadrowski K, Meyer ST, Mommer L, Weigelt A. From pots to plots: hier- archical trait-based prediction of plant performance in a mesic grassland. Journal of Ecology. 2016; 104(1):206–18. 91. Olofsson J, Oksanen L, Callaghan T, Hulme PE, Oksanen T, Suominen O. Herbivores inhibit climate- driven shrub expansion on the tundra. Global Change Biology. 2009; 15(11):2681–93. 92. Eskelinen A, Kaarleja¨ rvi E, Olofsson J. Herbivory and nutrient limitation protect warming tundra from lowland species’ invasion and diversity loss. Global Change Biology. 2017; 23(1):245–55. https://doi. org/10.1111/gcb.13397 PMID: 27343482 93. Speed JD, Austrheim G, Hester AJ, Mysterud A. Experimental evidence for herbivore limitation of the treeline. Ecology. 2010; 91(11):3414–20. https://doi.org/10.1890/09-2300.1 PMID: 21141202 94. Klein JA, Harte J, Zhao X-Q. Experimental warming causes large and rapid species loss, dampened by simulated grazing, on the Tibetan Plateau. Ecology Letters. 2004; 7(12):1170–9. 95. Post E, Pedersen C. Opposing plant community responses to warming with and without herbivores. Proceedings of the National Academy of Sciences. 2008; 105(34):12353–8. 96. Kaarleja¨ rvi E, Eskelinen A, Olofsson J. Herbivores rescue diversity in warming tundra by modulating trait-dependent species losses and gains. Nature Communications. 2017; 8(1):419. https://doi.org/10. 1038/s41467-017-00554-z PMID: 28871154 97. Harrison PA, Dunford RW, Holman IP, Cojocaru G, Madsen MS, Chen P-Y, et al. Differences between

low-end and high-end climate change impacts in Europe across multiple sectors. Regional Environ- mental Change. 2018; 19(3):695–709. 98. Kafle D, Wurst S, Dam N. Legacy effects of herbivory enhance performance and resistance of progeny plants. Journal of Ecology. 2018; 107(1):58–68. 99. Eskelinen A, Oksanen J. Changes in the abundance, composition and species richness of mountain vegetation in relation to summer grazing by reindeer. Journal of Vegetation Science. 2006; 17(2):245– 54. 100. Siefert A, Ritchie ME. Intraspecific trait variation drives functional responses of old-field plant commu- nities to nutrient enrichment. Oecologia. 2016; 181(1):245–55. https://doi.org/10.1007/s00442-016- 3563-z PMID: 26826004 101. Rozendaal DMA, Hurtado VH, Poorter L. Plasticity in leaf traits of 38 tropical tree species in response to light; relationships with light demand and adult stature. Functional Ecology. 2006; 20(2):207–16. PLOS ONE Transplant performance and trait responses to abiotic and biotic changes PLOS ONE | https://doi.org/10.1371/journal.pone.0276789 November 8, 2022 16 / 16