Vegetation and vertebrate abundance as drivers of bioturbation patterns along a climate gradient

37. Fick SE, Hijmans RJ. WorldClim 2: new 1-km spatial resolution climate surfaces for global land areas. Int J Climatol. 2017; 37: 4302–4315. https://doi.org/10.1002/joc.5086 38. U¨ bernickel K, Ehlers TA, Ershadi MR, Paulino L, Fuentes Espoz J-P, Maldonado A, et al. Time series of meteorological station data in the EarthShape study areas in the Coastal Cordillera, Chile. 2020 [cited 2 Nov 2021]. Available: https://dataservices.gfz-potsdam.de/panmetaworks/showshort.php?id= d4691c94-1d01-11eb-9603-497c92695674 39. Gabet EJ. Gopher bioturbation: Field evidence for non-linear hillslope diffusion. Earth Surf Process Landforms. 2000; 25: 1419–1428. https://doi.org/10.1002/1096-9837(200012)25:13<1419::AID- ESP148>3.0.CO;2–1 40. Richards PJ, Humphreys GS, Tomkins KM, Shakesby RA, Doerr SH. Bioturbation on wildfire-affected southeast Australian hillslopes: Spatial and temporal variation. Catena. 2011; 87: 20–30. https://doi.org/ 10.1016/j.catena.2011.05.003 41. Grigusova P, Larsen A, Achilles S, Klug A, Fischer R, Kraus D, et al. Area-wide prediction of vertebrate and invertebrate hole density and depth across a climate gradient in chile based on uav and machine learning. Drones. 2021; 5: 86. https://doi.org/10.3390/drones5030086 42. Horn BKP. Hill Shading and the Reflectance Map. Proc IEEE. 1981; 69: 14–47. https://doi.org/10.1109/ PROC.1981.11918 43. Ku¨ gler M, Hoffmann TO, Beer AR, U¨ bernickel K, Ehlers TA, Scherler D, Eichel J. (LiDAR) 3D Point Clouds and Topographic Data from the Chilean Coastal Cordillera. [cited 26 Jan 2022]. Available:

https://dataservices.gfz-potsdam.de/panmetaworks/showshort.php?id=4a3ddd07-7224-11ec-958d- ed2b0fcbcc9b 44. Huxley J. Problems Of Relative Growth: Huxley, Julian S.: Free Download, Borrow, and Streaming: Internet Archive. [cited 2 Nov 2021]. Available: https://archive.org/details/problemsofrelati033234mbp/ page/n10/mode/2up 45. Harper W V. Reduced Major Axis Regression: Teaching Alternatives to Least Reduced Major Axis Regression: Teaching Alternatives to Least Squares Squares. 2014 [cited 2 Nov 2021]. Available: https://digitalcommons.otterbein.edu/math_fac/24 46. Legendre. Package “lmodel2” Type Package Title Model II Regression. 2018. 47. Voeten CC. Using “buildmer” to automatically find & compare maximal (mixed) models. 2019. 48. Bates D, Maechler M, Bolker B. Package “lme4.” 2021. 49. Zhang D. Rsq: R-Squared and Related Measures. R package. 2020. 50. Chambers JM, Hastie TJ. Statistical Models. Stat Model S. 2017; 13–44. https://doi.org/10.1201/ 9780203738535-2 51. White CR. The allometry of burrow geometry. J Zool. 2005; 265: 395–403. https://doi.org/10.1017/ S0952836905006473 52. Milne T, Bull CM. Burrow choice by individuals of different sizes in the endangered pygmy blue tongue lizard Tiliqua adelaidensis. Biol Conserv. 2000; 95: 295–301. https://doi.org/10.1016/S0006-3207(00) 00040-9 53. Vleck D. Burrow structure and foraging costs in the fossorial rodent, Thomomys bottae. Oecologia 1981 493. 1981; 49: 391–396. https://doi.org/10.1007/BF00347605 PMID: 28310003

54. Peters RH, Wassenberg K. The effect of body size on animal abundance. Oecologia 1983 601. 1983; 60: 89–96. https://doi.org/10.1007/BF00379325 PMID: 28310539 55. Platt BF, Kolb DJ, Kunhardt CG, Milo SP, New LG. Burrowing through the literature: The impact of soil- disturbing vertebrates on physical and chemical properties of soil. Soil Sci. 2016; 181: 175–191. https:// doi.org/10.1097/SS.0000000000000150 56. Newsome AE, Corbett LK. Outbreaks of Rodents in Semi-Arid and Arid Australia: Causes, Preventions, and Evolutionary Considerations. 1975; 117–153. https://doi.org/10.1007/978-94-010-1944-6_6 57. Roper TJ. Badger Meles meles setts–architecture, internal environment and function. Mamm Rev. 1992; 22: 43–53. https://doi.org/10.1111/J.1365-2907.1992.TB00118.X 58. Abaturov B. D. The role of burrowing animals in the transport of mineral substances in the soil. Pedobio- logia (Jena). 1972; 12: 261–266. Available: https://ci.nii.ac.jp/naid/10030967703 59. Ellison L. The Pocket Gopher in Relation to Soil Erosion on Moutain Range. Ecology. 1946; 27: 101– 114. https://doi.org/10.2307/1932505 60. Humphreys GS. Earthen structures built by nymphs of the cicada Cyclochila australasiae (Donovan) (Homoptera: Cicadidae). Aust ent Mag. 1989; 16: 99–108. Available: https://search.informit.org/doi/ abs/10.3316/informit.118491194646625 PLOS ONE Vegetation and vertebrate abundance as drivers of bioturbation patterns along a climate gradient PLOS ONE | https://doi.org/10.1371/journal.pone.0264408 March 4, 2022 13 / 14