Interacting effects of habitat structure and seeding with oysters on the intertidal biodiversity of seawalls

33. Palomo M, People J, Chapman M, Underwood A. Separating the effects of physical and biological aspects of mussel beds on their associated assemblages. Marine Ecology Progress Series. 2007; 344:131–42. 34. Summerhayes SA, Bishop MJ, Leigh A, Kelaher BP. Effects of oyster death and shell disarticulation on associated communities of epibiota. Journal of Experimental Marine Biology and Ecology. 2009; 379(1– 2):60–7. 35. Andre´ C, Jonsson P, Lindegarth M. Predation on settling bivalve larvae by benthic suspension feeders: the role of hydrodynamics and larval behaviour. Marine Ecology Progress Series. 1993:183–92. 36. Commito JA, Boncavage EM. Suspension-feeders and coexisting infauna: an enhancement counterex- ample. Journal of Experimental Marine Biology and Ecology. 1989; 125(1):33–42. 37. Gratwicke B, Speight MR. Effects of habitat complexity on Caribbean marine fish assemblages. Marine Ecology Progress Series. 2005; 292:301–10. 38. Morris RL, Chapman MG, Firth LB, Coleman RA. Increasing habitat complexity on seawalls: Investigat- ing large-and small-scale effects on fish assemblages. Ecology and Evolution. 2017; 7(22):9567–79. https://doi.org/10.1002/ece3.3475 PMID: 29187990 39. Ushiama S, Mayer-Pinto M, Bugnot A, Johnston E, Dafforn K. Eco-engineering increases habitat avail- ability and utilisation of seawalls by fish. Ecological Engineering. 2019; 138:403–11. 40. Ferse SCA. Artificial reef structures and coral transplantation: fish community responses and effects on

coral recruitment in North Sulawesi/Indonesia: Dissertation. University of Bremen, Bremen, Germany; 2008. 41. Gregalis KC, Johnson MW, Powers SP. Restored oyster reef location and design affect responses of resident and transient fish, crab, and shellfish species in Mobile Bay, Alabama. Transactions of the American Fisheries Society. 2009; 138(2):314–27. 42. Ng CSL, Lim SC, Ong JY, Teo LMS, Chou LM, Chua KE, et al. Enhancing the biodiversity of coastal defence structures: transplantation of nursery-reared reef biota onto intertidal seawalls. Ecological Engineering. 2015; 82:480–6. 43. Scanes E, Johnston EL, Cole VJ, O’Connor WA, Parker LM, Ross PM. Quantifying abundance and dis- tribution of native and invasive oysters in an urbanised estuary. Aquatic Invasions. 2016; 11(4):425–36. 44. Kohler KE, Gill SM. Coral Point Count with Excel extensions (CPCe): A Visual Basic program for the determination of coral and substrate coverage using random point count methodology. Computers and Geosciences. 2006; 32(9):1259–69. 45. Cappo MM, Harvey EE, Shortis MM, editors. Counting and measuring fish with baited video tech- niques-an overview. Proceedings of the Australian Society for Fish Biology Workshop, Hobart, August 2007-pages: 101–114; 2007. 46. R Core Team. R: A Language and Environment for Statistical Computing. Vienna, Austria: R Founda- tion for Statistical Computing; 2016. 47. Wickham H. ggplot2. Computational Statistics. 2011; 3(2):180–5. 48. McAfee D O’connor WA, Bishop MJ. Fast-growing oysters show reduced capacity to provide a thermal

refuge to intertidal biodiversity at high temperatures. Journal of Animal Ecology. 2017; 86(6):1352–62. https://doi.org/10.1111/1365-2656.12757 PMID: 28913869 49. Bishop MJ, Mayer-Pinto M, Airoldi L, Firth LB, Morris RL, Loke LHL, et al. Effects of ocean sprawl on ecological connectivity: impacts and solutions. Journal of Experimental Marine Biology and Ecology. 2017. 50. Archambault P, Bourget E. Scales of coastal heterogeneity and benthic intertidal species richness, diversity and abundance. Marine Ecology Progress Series. 1996; 136(1):111–21. 51. McQuaid CD, Porri F, Nicastro K, Zardi G. Simple, scale-dependent patterns emerge from very complex effects: an example from the intertidal mussels Mytilus galloprovincialis and Perna perna. Oceanogra- phy and Marine Biology: An Annual Review. 2015; 53:127–56. 52. MacArthur M, Naylor LA, Hansom JD, Burrows MT, Loke LH, Boyd I. Maximising the ecological value of hard coastal structures using textured formliners. Journal of Ecological Engineering. 2019; 1:100002. 53. Bateman D, Bishop MJ. The environmental context and traits of habitat-forming bivalves influence the magnitude of their ecosystem engineering. Marine Ecology Progress Series. 2016; 563:95–110. 54. Grabowski JH, Peterson CH. Restoring oyster reefs to recover ecosystem services. Ecosystems engi- neers: plants to protists. 42007. p. 281–98. 55. Wilkie EM, Bishop MJ, O’Connor WA. Are native Saccostrea glomerata and invasive Crassostrea gigas oysters’ habitat equivalents for epibenthic communities in south-eastern Australia? Journal of Experi-

mental Marine Biology and Ecology. 2012; 420:16–25. PLOS ONE Effects of microhabitats and seeding with oysters on intertidal biodiversity PLOS ONE | https://doi.org/10.1371/journal.pone.0230807 July 16, 2020 20 / 21