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Does increasing biodiversity in an urban woodland setting promote positive emotional responses in humans? A stress recovery experiment using 360-degree videos of an urban woodland

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Acknowledgments The authors would like to thank Ms Liwen Zhang and Mr John Land for their comments on the manuscript. Author Contributions Conceptualization: Simone Farris, Nicola Dempsey, Kirsten McEwan, Helen Hoyle, Ross Cameron. Data curation: Simone Farris. Formal analysis: Simone Farris, Nicola Dempsey, Ross Cameron. Investigation: Simone Farris, Ross Cameron. Methodology: Simone Farris, Nicola Dempsey, Kirsten McEwan, Ross Cameron. Supervision: Nicola Dempsey, Kirsten McEwan, Ross Cameron. Writing – original draft: Simone Farris, Ross Cameron. Writing – review & editing: Simone Farris, Nicola Dempsey, Kirsten McEwan, Helen Hoyle, Ross Cameron. References 1. Van Den Berg M, Van Poppel M, Van Kamp I, Andrusaityte S, Balseviciene B, Cirach M, et al. Visiting green space is associated with mental health and vitality: A cross-sectional study in four european cit- ies. Health Place. 2016; 38: 8–15. https://doi.org/10.1016/j.healthplace.2016.01.003 PMID: 26796323 2. Wood L, Hooper P, Foster S, Bull F. Public green spaces and positive mental health–investigating the relationship between access, quantity and types of parks and mental wellbeing. Health Place. 2017; 48: 63–71. https://doi.org/10.1016/j.healthplace.2017.09.002 PMID: 28942343 3. Vanaken G-J, Danckaerts M. Impact of Green Space Exposure on Children’s and Adolescents’ Mental Health: A Systematic Review. Int J Environ Res Public Health. 2018; 15. https://doi.org/10.3390/ ijerph15122668 PMID: 30486416 4. Frumkin H, Bratman G, Breslow S, Cochran B, Kahn P, Lawler J, et al. Nature Contact and Human

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Health: A Research Agenda. Environ Health Perspect Online. 2017; 125. https://doi.org/10.1289/ EHP1663 PMID: 28796634 5. Kondo MC, Jacoby SF, South EC. Does spending time outdoors reduce stress? A review of real-time stress response to outdoor environments. Health Place. 2018; 51: 136–150. https://doi.org/10.1016/j. healthplace.2018.03.001 PMID: 29604546 6. Markevych I, Schoierer J, Hartig T, Chudnovsky A, Hystad P, Dzhambov AM, et al. Exploring pathways linking greenspace to health: Theoretical and methodological guidance. Environ Res. 2017; 158. https://doi.org/10.1016/j.envres.2017.06.028 PMID: 28672128 7. Yao W, Zhang X, Gong Q. The effect of exposure to the natural environment on stress reduction: A meta-analysis. Urban For Urban Green. 2021; 57: 1–12. https://doi.org/10.1016/j.ufug.2020.126932 8. Kaplan S. The restorative benefits of nature: Toward an integrative framework. J Environ Psychol. 1995; 15: 169–182. https://doi.org/10.1016/0272-4944(95)90001-2 9. Ulrich RS, Simons RF, Losito BD, Fiorito E, Miles MA, Zelson M. Stress recovery during exposure to natural and urban environments. J Environ Psychol. 1991; 11: 201–230. https://doi.org/10.1016/S0272- 4944(05)80184-7 10. Kellert SR, Wilson EO. The Biophilia hypothesis. Kellert SR, Wilson EO, editors. Washington, D.C.: Washington, D.C.: Island Press, 1993; 1993. 11. Li Q, Kobayashi M, Wakayama Y, Inagaki H, Katsumata M, Hirata Y, et al. Effect of Phytoncide from Trees on Human Natural Killer Cell Function. Int J Immunopathol Pharmacol. 2009; 22: 951–959.

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https://doi.org/10.1177/039463200902200410 PMID: 20074458 12. Haahtela T. A biodiversity hypothesis. Allergy. 2019; 74: 1445–1456. https://doi.org/10.1111/all.13763 PMID: 30835837 PLOS ONE Does increasing biodiversity in an urban woodland setting, promote positive emotional responses in humans? PLOS ONE | https://doi.org/10.1371/journal.pone.0297179 February 7, 2024 16 / 20