Biodiversity of ecosystems in an arid setting: The late Albian plant communities and associated biota from eastern Iberia

Table 1. (Continued) Taxon Section Known or probable parent affinity Possible preferred environment Hammenia fredericksburgensis SJ, CA, AP Antophyta; Chloranthaceae Ground cover/hygrophytic (river banks, pioneer) Impardecispora cf. apiverrucata CA Monilophyta; Lygodiaceae Ground cover/hygrophytic (understory, wet lowland) Impletosphaeridium spp. SJ, CA, AP Dinoflagellata: uncertain affinity Marine Inaperturopollenites dubius SJ, CA, AP Gymnospermophyta; Coniferales; Cupressaceae (taxodioids) Canopy/hygrophytic (mire, marshes, river shores, wet lowland) Inaperturopollenites spp. SJ, CA, AP Gymnospermophyta; Coniferales; Araucariaceae?, Cupressaceae? Canopy/xerophytic? (lowland, coastal?) Ischyosporites crateris CA Monilophyta; Schizaeales; Schizaeaceae Ground cover/hygrophytic (understory, wet lowland) Ischyosporites cf. pseudoreticulatus AP Ischyosporites cf. punctatus CA Ischyosporites variegatus CA Ischyosporites sp. SJ, CA Jusinghipollis cf. ticoensis SJ, CA, AP Antophyta; uncertain affinity Ground cover?/hygrophytic?/canopy? ?Kallosphaeridium spp SJ Dinoflagellata: Gonyaulacaceae Marine Kiokansium unituberculatum CA, AP Dinoflagellata: uncertain affinity Marine Kraeuselisporites sp. (= Patellasporites? echinatus) SJ Lycophyta; Selaginellaceae? Ground cover, xerophytic? Kraeuselisporites spp. CA, AP Laevigatosporites ovatus CA, AP Monilophyta; uncertain affinities Ground cover/hygrophytic (lowland) Laevigatosporites spp. SJ, CA Leptolepidites cf. crassibalteus CA Unknown—Lycophyta?, Monilophyta? Ground cover/hygrophytic?

Leptolepidites proxigranulatus CA cf. Leptolepidites sp. SJ Leptolepidites spp. SJ, CA, AP Liliacidites cf. clavatus CA, AP Antophyta; probable monocot Ground cover/hygrophytic (wet lowland, marshes) Liliacidites cf. kiowaensis CA Liliacidites tectatus CA, AP Liliacidites spp. SJ, CA, AP “Liliacidites” minutus SJ, CA, AP Antophyta; Laurales Canopy?/xerophytic? (lowland) Lophotriletes sp. AP Monilophyta; Botryopteridales Ground cover/hygrophytic (wet lowland) ?Lusatisporites dettmannae SJ, CA, AP Monilophyta; uncertain affinities Ground cover/hygrophytic? Lycopodiumsporites sp. AP Lycophyta; Lycopodiaceae Ground cover/hygrophytic (lowland, mires, wet environments) cf. Margocolporites sp. SJ, CA Antophyta; Eudicots Canopy?/xerophytic? (lowland) Matonisporites crassiangulatus CA Monilophyta; Matoniaceae Ground cover/xerophytic? (lowland) Matonisporites sp. AP Micrhystridium sp. SJ, CA, AP Acritarcha: Acanthomorphitae Marine Microfoveolatosporis pseudoreticulatus CA Monilophyta; uncertain affinities Ground cover/hygrophytic Mirovia gothanii CA, AP Gymnospermophyta; Coniferales; Miroviaceae Canopy/xerophytic (lowland, marshes) Monocolpopollenites sp. SJ, CA, AP Antophyta; Arecales, Arecaceae Canopy/xerophytic? (lowland) (Continued) PLOS ONE Late Albian desert biota from Iberia PLOS ONE | https://doi.org/10.1371/journal.pone.0282178 March 2, 2023 15 / 51