Biodiversity of ecosystems in an arid setting: The late Albian plant communities and associated biota from eastern Iberia

Table 1. (Continued) Taxon Section Known or probable parent affinity Possible preferred environment Crybelosporites pannuceus SJ, CA, AP Monilophyta; Marsileaceae Ground cover/hygrophytic Cupuliferoidaepollenites parvulus CA Antophyta; non-magnoliid dicotyledon Canopy?/xerophytic? (lowland) Cupuliferoidaepollenites sp. SJ, CA, AP Cyathidites australis SJ, CA, AP Monilophyta; uncertain affinities Ground cover/hygrophytic (understory, wet lowland) Cyathidites minor SJ, CA, AP Cycadopites carpentieri CA Gymnospermophyta; uncertain affinity (Bennettitales, Cycadales, Czekanowskiales, Ginkgoales, Gnetales, Pentoxylales) Canopy/xerophytic (lowland, coastal) Cycadopites follicularis SJ, CA, AP Cycadopites spp. SJ, CA, AP Cyclonephelium chabacca CA, AP Dinoflagellata: Areoligeraceae Marine Cyclonephelium vannophorum CA Cyclonephelium spp. SJ, CA, AP Cymathiosphaera spp. CA, AP Chlorophyta (Prasinophyte) Marine Dammarites cf. albens AP Gymnospermophyta; Coniferales; Araucariaceae Canopy/xerophytic (coastal) Dammarites spp. SJ Dapsilidinium sp. SJ Dinoflagellata: uncertain affinity Marine Deltoidospora spp. SJ, CA, AP Monilophyta; uncertain affinities Ground cover/hygrophytic (understory, wet lowland) Densoisporites velatus CA Lycophyta; Pleuromeiaceae? Ground cover/hygrophytic (coastal, mires) Densoisporites sp. SJ, AP Dichastopollenites dunveganensis SJ, CA Antophyta; uncertain affinity Ground cover?, floating plant?/hygrophytic Dichastopollenites ghazalatensis CA Dichastopollenites cf. reticulatus AP Dichastopollenites spp. SJ, CA, AP

Dictyophyllidites equiexinus CA, AP Monilophyta; Dipteridaceae/Matoniaceae Ground cover/hygrophytic? Dictyophyllidites harrisii SJ, CA, AP Dictyophyllidites sp. CA Distachyapites sp. SJ Gymnospermophyta; Gnetales; Ephedraceae Shrubs?/xerophytic (lowland, coastal, upland) Distaltriangulisporites irregularis CA Monilophyta; Schizaeales; Schizaeaceae Ground cover/hygrophytic (understory, wet lowland) Distaltriangulisporites maximus CA Distaltriangulisporites cf. maximus CA Distaltriangulisporites perplexus SJ Distaltriangulisporites cf. perplexus CA Distaltriangulisporites sp. CA Distaverrusporites sp. AP Monilophyta; uncertain affinities Ground cover/hygrophytic? Doyleipollenites robbinsiae SJ, AP Antophyta; uncertain affinity Ground cover?/hygrophytic? Dryadopollis vestalis CA, AP Antophyta; non-magnoliid dicotyledon Ground cover?, shrubs?/xerophytic? (lowland) Echimonocolpites sp. AP Antophyta; uncertain affinity (Nymphaeaceae?, Monocots?) Floating plants/hygrohytic (mire, marshes) Echinatisporis varispinosus SJ Lycophyta; Selaginellaceae Ground cover, xerophytic? Echinatisporis sp. CA, AP Echinipollis cenomanensis CA Antophyta; uncertain affinity Ground cover?/xerophytic? (Continued) PLOS ONE Late Albian desert biota from Iberia PLOS ONE | https://doi.org/10.1371/journal.pone.0282178 March 2, 2023 13 / 51