Fifty years after: A taxonomic revision of the amphibian species from the Ecuadorian biodiversity hotspot Abra de Zamora, with description of two new Pristimantis species

supratympanic fold present; (3) snout subacuminate in dorsal view, rounded in profile; can- thus rostralis straight in dorsal view, rounded in profile; (4) upper eyelid bearing 1 to 3 larger tubercles (trait more visible in life), about 84% IOD; cranial crests absent but prominent) (or > < -shaped scapular folds are present (trait more visible in life); (5) dentigerous processes of vomers prominent, oblique, ovoid, separated medially by distance equal or lower than the width of processes; each processes bearing 3 to 5 teeth; (6) males lacking vocal sac and slits; nuptial pads absent; (7) Finger I shorter than Finger II; discs on fingers broadly expanded, truncated; circumferential grooves present; fingers bearing narrow lateral fringes; (8) subarti- cular tubercles prominent; supernumerary palmar tubercles evident; palmar tubercle partially (sometimes completely) divided into a larger (inner) and a smaller (outer) tubercles; thenar tubercle elliptical, smaller than the palmar tubercle but larger than either palmar tubercles (inner or outer); (9) prominent ulnar tubercles present (trait more visible in life); (10) heel with one large, conical and several smaller tubercles; outer edge of tarsus bearing a row of enlarged spine like tubercles (trait more visible in life); inner tarsal tubercles coalesced into a short tarsal fold; (11) inner metatarsal tubercle elongated, not compressed, about 4 to 6x ovoid, subconical (in profile), outer metatarsal tubercle; supernumerary plantar tubercles pres-

ent; (12) Toe V much longer than Toe III; discs on toes broadly expanded, truncated, slightly smaller than those of fingers; circumferential grooves present; toes bearing narrow lateral Fig 22. Color variation of Pristimantis cryptomelas in life. A. female: MUTPL 168; B. male: MUTPL 470; C. subadult: MUTPL 135; D. juvenile: MUTPL 493. Abra de Zamora (B, C), Bosque Protector Washapamba (A), Parque Nacional Podocarpus—Cajanuma (D). Most common coloration of this species is presented in A. https://doi.org/10.1371/journal.pone.0238306.g022 PLOS ONE Taxonomic revision of the amphibian species from Abra de Zamora PLOS ONE | https://doi.org/10.1371/journal.pone.0238306 September 10, 2020 39 / 59