Fifty years after: A taxonomic revision of the amphibian species from the Ecuadorian biodiversity hotspot Abra de Zamora, with description of two new Pristimantis species

to be light bronze and not reddish bronze), and we add descriptions of the characters that are more evident in life. The diagnosis provided herein is based on 14 specimens from the original description (KU 142151–64), two adult females (MUTPL 311, 388) one adult male (MUTPL 675) and two juveniles (MUTPL 732, MZUA 2491) collected from the type locality. Pristimantis colodactylus is part of the P. colodactylus subgroup, and as such it is closely related to P. matildae sp. nov., P. muranunka and an additional undescribed species from Reserva Tapichalaca (Fig 7). The genetic distance between P. colodactylus and its closest rela- tives from the subgroup ranges between 5.4–7.5% (S3 Table). Pristimantis matildae sp. nov. Sze´kely, Eguiguren, Ordo´ñez-Delgado, Armijos-Ojeda, and Sze´kely urn:lsid:zoobank.org:act:E2E120CF-C257-4A95-9052-879CA1387C89 (Figs 16–19) Common English name. Matilde’s Rain Frog Common Spanish name. Cutı´n de Matilde Etymology. The specific epithet is a noun in the genitive case and is a patronym for Matilde Hidalgo Navarro (1889–1974). She was the first woman to obtain a medical degree in the country, and the first woman in Ecuador (and Latin America) to exercise the right to vote in a Fig 16. Holotype of Pristimantis matildae sp. nov. (MUTPL 731, adult male), SVL 20.2 mm, in life. A. Lateral view; B. Ventral view; C. Dorsal view. https://doi.org/10.1371/journal.pone.0238306.g016 PLOS ONE Taxonomic revision of the amphibian species from Abra de Zamora PLOS ONE | https://doi.org/10.1371/journal.pone.0238306 September 10, 2020 31 / 59