Fifty years after: A taxonomic revision of the amphibian species from the Ecuadorian biodiversity hotspot Abra de Zamora, with description of two new Pristimantis species

females and 72% IOD in males; cranial crests absent; (5) dentigerous processes of vomers con- cealed in buccal mucosa; each processes bearing 2 to 6 teeth; (6) males lacking vocal sac and slits; (7) fingers very short and stocky; Finger I shorter than Finger II; discs on fingers slightly expanded, rounded; circumferential grooves present; fingers bearing lateral fringes; (8) subar- ticular tubercles prominent; supernumerary palmar tubercles numerous, continuing onto fin- gers; palmar tubercle completely divided into a larger (inner) and a smaller (outer) tubercles; thenar tubercle oval, the same size or slightly smaller than the inner palmar tubercle; (9) ulnar tubercles absent; (10) heel bearing one moderate-sized conical tubercle and several small tubercles (trait more visible in life); inner and outer edges of tarsus lacking tubercles or folds; (11) inner metatarsal tubercle broadly ovoid, about 4 to 5x rounded outer metatarsal tubercle; supernumerary plantar tubercles numerous, continuing onto toes; (12) toes very short and stocky; Toe V much longer than Toe III; discs on toes slightly expanded, rounded, about same size as those on fingers; circumferential grooves present; toes bearing prominent lateral fringes; webbing basal; (13) in life, dorsum various shades of brown with or without pale dor- solateral stripes, interorbital bars, or dark X-shaped markings on the back; venter cream with Fig 15. Color variation of Pristimantis colodactylus in life. A–B females: A. MUTPL 311, B. MUTPL 388; C. male:

MUTPL 675; D–E juveniles: D. MZUA 2491, E. MUTPL 732. All specimens are from Abra de Zamora. https://doi.org/10.1371/journal.pone.0238306.g015 PLOS ONE Taxonomic revision of the amphibian species from Abra de Zamora PLOS ONE | https://doi.org/10.1371/journal.pone.0238306 September 10, 2020 29 / 59